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Studies have revealed that MSCs secrete several soluble factors that contribute to their immunomodulatory effects and regulate immune cell responses [41, 42]

These genes were then compared to genes belonging to the leading edge of the same gene signature from i4F MEFs treated with doxycycline in vitro for 72 h, as compared to OSKM MEFs treated with vitamin B 12 (that is, genes in MsigDB M13537 whose upregulation was relieved by B 12 supplementation in vitro)

Because they're missing certain amino, so you got to combine them just right

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This is why a thorough screening process is so vital

This is known as functional B12 deficiency