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Moreover, the expression of key spermatogenesis markers, including UCHL1, PLZF, SCP3, and PCNA, which were reduced in aged testes, were significantly restored or increased following FOXO4-DRI treatment

Tragically this next audit published in 1985 only two years later, makes things a whole lot worse

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Conservation of Buck's Spermatozoa during Cooling Storage Period through Cooling Medium Supplementation with L-Carnitine

These likely include the matrix metalloproteases MMP2 and MMP9, whose expression upregulates following LEC injury, potentially leading to cleavage of the LAP protein to release active TGF (49, 173, 227, 228)

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